Tardigrade Tools
CHECK LIST OF TARDIGRADE TAXA
(download the most updated list of the tardigrade taxa)
TAXONOMIC KEYS
(the following articles present the most updated taxonomic keys for several tardigrade taxa)
Heterotardigrada
Arthrotardigrada
Arthrotardigrada genera: Fontoura, P., Bartels, P. J., Jørgensen, A., Kristensen, R. M., & Hansen, J. G. (2017). A dichotomous key to the genera of the Marine Heterotardigrades (Tardigrada). Zootaxa, 4294(1), 1-45.
Archechiniscidae
Archechiniscus species: Bartels, P. J., Fontoura, P., & Nelson, D. R. (2018). Marine tardigrades of the Bahamas with the description of two new species and updated keys to the species of Anisonyches and Archechiniscus. Zootaxa, 4420(1), 43-70.
Batillipedidae
Batillipes species:Santos E., Rubal M., Veiga P., Rocha C. & Fontoura P. (2018). Batillipes (Tardigrada, Arthrotardigrada) from the Portuguese coast with the description of two new species and a new dichotomous key for all species. European Journal of Taxonomy, (425), 1-32.
Coronarctidae
Coronarctus species: Gomes-Júnior, E., Santos, É., da Rocha, C. M., Santos, P. J., & Fontoura, P. (2020). The deep-sea genus Coronarctus (Tardigrada, Arthrotardigrada) in Brazil, south-western Atlantic Ocean, with the description of three new species. Diversity, 12(2), 63.
Stygarctidae
Megastygarctides species: Ürkmez, D., Ostrowska, M., Roszkowska, M., Gawlak, M., Zawierucha, K., Kristensen, R. M., & Kaczmarek, Ł. (2018). Description of Megastygarctides sezginii sp. nov.(Tardigrada: Arthrotardigrada: Stygarctidae) from the Turkish Black Sea coast and a key to the genus Megastygarctides. Marine Biology Research, 14(1), 1-16.
Styraconyxidae
Angursa species: Fujimoto, S., & Hansen, J. G. (2019). Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan. European Journal of Taxonomy, (510).
Tanarctidae
Tanarctus species: Fujimoto, S., Miyazaki, K., & Suzuki, A. C. (2013). A new marine tardigrade, Tanarctus diplocerus (Arthrotardigrada: Halechiniscidae) from Japan. Journal of the Marine Biological Association of the United Kingdom, 93(4), 955-961.
Echiniscoidea
Echiniscoididae
Echiniscoididae: Gąsiorek, P. & Kristensen R. M. (2022). New marine heterotardigrades lineages from the tropics. The European Zoological Journal. 89:1, 719-754.
Anisonyches species: Bartels, P. J., Fontoura, P., & Nelson, D. R. (2018). Marine tardigrades of the Bahamas with the description of two new species and updated keys to the species of Anisonyches and Archechiniscus. Zootaxa, 4420(1), 43-70.
Echiniscidae
Bryodelphax species: Fontoura, P., Pilato, G., & Lisi, O. (2008). Echiniscidae (Tardigrada, Heterotardigrada) from Faial and Pico Islands, the Azores, with the description of two new species. Zootaxa, 1693, 49-61.
Cornechiniscus species: Gąsiorek, P. & Michalczyk, Ł. (2020). Revised Cornechiniscus (Heterotardigrada) and new phylogenetic analyses negate echiniscid subfamilies and tribes. Royal Society Open Science, 7: 200581.
Echiniscus bigranulatus group: Michalczyk, L. & Kaczmarek, L. (2007). Echiniscus ganczareki, a new species of Tardigrada (Heterotardigrada: Echiniscidae, bigranulatus group) from Costa Rica. Zootaxa, 1471, 1525.
Mopsechiniscus species: Dastych, H. (2001). Notes on the Revision of the Genus Mopsechiniscus (Tardigrada). Zoologisher Anzeiger, 240, 299308.
Hypechiniscus species: Gasiorek et al. (2021). New Asian and Nearctic Hypechiniscus species (Heterotardigrada: Echiniscidae) signalize a pseudocryptic horn of plenty. Zoological Journal of the Linnean Society, 192, 794-852.
Eutardigrada
Eutardigrada genera: Pilato, G., & Binda, M. G. (2010). Definition of families, subfamilies, genera and subgenera of the Eutardigrada, and keys to their identification. Zootaxa, 2404, 1-54.
Eutardigrada genera: Nelson, D. R., Guidetti, R., & Rebecchi, L. (2015). Phylum Tardigrada. In Thorp, J. H., & Covich, A. P. (Eds.). Ecology and classification of North American freshwater invertebrates. Academic press.. pp. 347-380.
Eutardigrada genera: Bingemer, J., & Hohberg, K. (2017). An illustrated identification key to the eutardigrade species (Tardigrada, Eutardigrada) presently known from European soils. Soil Organisms 89(3): 127-149.
Apochela
Milnesium species: Morek, W., Gąsiorek, P., Stec, D., Blagden, B. & Michalczyk, Ł. (2016). Experimental taxonomy exposes ontogenetic variability and elucidates the taxonomic value of claw configuration in Milnesium Doyère, 1840 (Tardigrada: Eutardigrada: Apochela). Contributions to Zoology, 85(2): 173-200.
Milnesium species of the granulatum group: Morek, W., Wałach, K., & Michalczyk, Ł. (2022) Rough backs: taxonomic value of epicuticular sculpturing in the genus Milnesium Doyère, 1840 (Tardigrada: Apochela), Scientific Reports 12: 9857. https://doi.org/10.1038/s41598-022-10758-z
Parachela
Eohypsibiidae
Bertolanius species: Hansen, J. G., Kristensen, R. M., Bertolani, R., & Guidetti, R. (2017). Comparative analyses of Bertolanius species (Eohypsibiidae; Eutardigrada) with the description of Bertolanius birnae sp. nov. from northern polar regions. Polar Biology, 40(1), 123-140.
Hypsibiidae
Diphascon species of pingue group: Fontoura, P., & Pilato, G. (2007). Diphascon (Diphascon) faialense sp. nov. a new species of Tardigrada (Eutardigrada, Hypsibiidae) from Azores and a key to the species of the D. pingue group. Zootaxa, 1589(4).
Hypsibius species of the dujardini group: Gąsiorek, P., Stec, D., Morek, W., & Michalczyk, Ł. (2018). An integrative redescription of Hypsibius dujardini (Doyère, 1840), the nominal taxon for Hypsibioidea (Tardigrada: Eutardigrada). Zootaxa, 4415(1), 45-75.
Itaquasconinae
Itaquasconinae genera: Gąsiorek, P., & Michalczyk, Ł. (2020). Phylogeny of Itaquasconinae in the light of the evolution of the flexible pharyngeal tube in Tardigrada. Zoologica Scripta, 49(4), 499-515.
Adropion & Diphascon without drop-like thickening: Gąsiorek, P., Blagden, B., Morek, W., & Michalczyk, Ł. (2023). What is a strongsynapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela). Zoological Journal of the Linnean Society, zlad151.
Guidettion species: Gąsiorek, P., Blagden, B., Morek, W., & Michalczyk, Ł. (2023). What is a strongsynapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela). Zoological Journal of the Linnean Society, zlad151.
Itaquascon species: Gąsiorek, P., Blagden, B., Morek, W., & Michalczyk, Ł. (2023). What is a strongsynapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela). Zoological Journal of the Linnean Society, zlad151
Platicrista species: Gąsiorek, P., Blagden, B., Morek, W., & Michalczyk, Ł. (2023). What is a strongsynapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela). Zoological Journal of the Linnean Society, zlad151
Raribius species: Gąsiorek, P., Blagden, B., Morek, W., & Michalczyk, Ł. (2023). What is a strongsynapomorphy? Redescriptions of Murrays type species and descriptions of new taxa challenge the systematics of Hypsibiidae (Eutardigrada: Parachela). Zoological Journal of the Linnean Society, zlad151
Arctodiphascon species: Tumanov, D. V., & Tsvetkova, A. Y. (2023). Some have drops and some do not, but can we rely on that? Re-investigation of Diphascon tenue (Tardigrada: Eutardigrada) with discussion of the phylogeny and taxonomy of the superfamily Hypsibioidea. Zoosystematica Rossica, 32, 50-74.
Ramazzottiidae
Ramazzottius species: Biserov, V. I. (1998). Tardigrades of the Caucasus with a taxonomic analysis of the genus Ramazzottius (Parachela: Hypsibiidae). Zoologisher Anzeiger, 236(1997), 139-159.
Calohypsibiidae
Calohypsibius species: Michalczyk, L., & Kaczmarek, L. (2005). The first record of the genus Calohypsibius Thulin, 1928 (Eutardigrada: Calohypsibiidae) from Chile (South America) with a description of a new species Calohypsibius maliki. New Zealand Journal of Zoology, 32(4), 287-292.
Isohypsibioidea
Isohypsibioidea: Gąsiorek, P., Stec, D., Morek, W., & Michalczyk, Ł. (2019). Deceptive conservatism of claws: distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contributions to Zoology, 88(1), 78-132.
Isohypsibiidae
Doryphoribius species: Pérez-Pech, W. A., Anguas-Escalante, A. B. R. I. L., Cutz-Pool, L. Q., & Guidetti, R. (2017). Doryphoribius chetumalensis sp. nov.(Eutardigrada: Isohypsibiidae) a new tardigrade species discovered in an unusual habitat of urban areas of Mexico. Zootaxa, 4344(2), 345-356.
Pseudobiotus species: Chang, Kaczmarek, Lee, Michalczyk (2007). Pseudobiotus spinifer, a New Tardigrade Species (Eutardigrada: Hypsibiidae) from Nakdong River, South Korea, with a Redescription of P. vladimiri Biserov, Dudichev & Biserova. Zoological Science 24: 623629.
Thulinius species: Kaczmarek, Bertolani, Nedzynska-Stygar (2010). Thulinius saltursus comb. nov.: a new systematic position for Isohypsibius saltursus Schuster, Toftner & Grigarick, 1978 (Eutardigrada: Hypsibiidae) and a key for the genus Thulinius. Zootaxa 2483: 5864.
Macrobiotidae
Calcarobiotus species: Kaczmarek, Ł., Michalczyk, Ł., & Guidetti, R. (2006). Description of the new species Calcarobiotus (C.) longinoi sp. nov.(Eutardigrada, Macrobiotidae) from Costa Rica with the diagnostic key to the genus Calcarobiotus. Italian Journal of Zoology, 73(3), 247-253.
Diaforobiotus species: Stec, D. (2023). Integrative taxonomy helps to revise systematics and questions the purported cosmopolitan nature of the type species within the genus Diaforobiotus (Eutardigrada: Richtersiusidae). Organisms Diversity & Evolution, 23(2), 309-328.
Macrobiotus species of hufelandi group: Kaczmarek, Ł., & Michalczyk, Ł. (2017). The Macrobiotus hufelandi group (Tardigrada) revisited. Zootaxa, 4363(1), 101.
Macrobiotus species of pseudohufelandi complex / Xerobiotus species: Vecchi et al. 2022. Macrobiotus naginae sp. nov., a new xerophilous tardigrade species from Rokua Sand Dunes (Finland). Zoological Studies 61:22. https://doi:10.6620/ZS.2022.61-22
Mesobiotus species. Stec, D. (2022). An Integrative Description of Two New Mesobiotus Species (Tardigrada: Eutardigrada: Macrobiotidae) with Updated Genus Phylogeny. Zoological Studies, 61.
Minibiotus species: Claxton, S. K. (1998). A revision of the genus Minibiotus (Tardigrada: Macrobiotidae) with descriptions of eleven new species from Australia. Records-Australian Museum, 50: 125-160.
Minibiotus species of sidereus group: Dueñas-Cedillo et al (2021). Species-Specific Cuticular Phenotypes in Eutardigrada: A Morphometric Approach to Analyze the Variation of Star-Shaped Pores in Minibiotus Species. Diversity, 13, 307.
Paramacrobiotus speces: Kaczmarek, Ł., Gawlak, M., Bartels, P. J., Nelson, D. R., & Roszkowska, M (2017). Revision of the genus Paramacrobiotus Guidetti et al., 2009 with the description of a new species, re-descriptions and a key. Annales Zoologici 67(4): 627-656.
Murrayidae
Murrayidae genera: Guidetti, R., Giovannini, I., Del Papa, V., Ekrem, T., Nelson, D. R., Rebecchi, L., & Cesari, M. (2022). Phylogeny of the asexual lineage Murrayidae (Macrobiotoidea, Eutardigrada) with the description of Paramurrayon gen. nov. and Paramurrayon meieri sp. nov. Invertebrate Systematics, 36(12), 1099-1117.
Murrayon species: Guidetti, R., Giovannini, I., Del Papa, V., Ekrem, T., Nelson, D. R., Rebecchi, L., & Cesari, M. (2022). Phylogeny of the asexual lineage Murrayidae (Macrobiotoidea, Eutardigrada) with the description of Paramurrayon gen. nov. and Paramurrayon meieri sp. nov. Invertebrate Systematics, 36(12), 1099-1117.
Paramurrayon species: Guidetti, R., Giovannini, I., Del Papa, V., Ekrem, T., Nelson, D. R., Rebecchi, L., & Cesari, M. (2022). Phylogeny of the asexual lineage Murrayidae (Macrobiotoidea, Eutardigrada) with the description of Paramurrayon gen. nov. and Paramurrayon meieri sp. nov. Invertebrate Systematics, 36(12), 1099-1117.
Dactylobiotus species : Kaczmarek, Ł., Schabetsberger, R., Litwin, M., & Michalczyk, Ł. (2012). A new freshwater eutardigrade from Fiji and Vanuatu (Oceania), with remarks on the genus Dactylobiotus. New Zealand Journal of Zoology, 39(4), 311-318.
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